... are able to support both specialist and more generalist fauna. The combination of soft sediment with hard substrate increases seabed heterogeneity, with dominance by one habitat type leading to greater homogeneity. According to the habitat heterogeneity hypothesis, more heterogeneous areas should support...

... for psychrophilic arthropods was assessed. The karst dolines revealed significant habitat heterogeneity, including warm plateaus and S-facing slopes and colder and wetter bottoms and N-facing slopes. The inner sites of the dolines had an overall higher abundance and species richness compared to the plateau sites...